The Anycloceratida were a diverse order of ammonites most closely related to the Lytoceratina . They evolved during the Late Jurassic but were not very common until the Cretaceous period, when they rapidly diversified and become one of the most distinctive components of Cretaceous marine faunas. They have been recorded from every continent and many are used as zonal or index fossils. The most distinctive feature of the majority of the Ancyloceratida is the tendency for most of them to have shells that are not regular spirals like most other ammonites. These irregularly-coiled ammonites are called heteromorph ammonites, in contrast to regularly-coiled ammonites, which are called homomorph ammonites.

In the more primitive forms, the shell departs only slightly from being a perfect spiral, with only the last, outermost whorl being open, forming a hook underneath the main spiral. In such forms the spiral was the chambered, buoyant part of the shell, and the hook was the living chamber in which the soft body of the ammonite resided. Examples of such types were Ancyloceras , Protanisoceras , and Tropaeum . The more advanced heteromorphs departed from such forms radically. The shells of Ptychoceras consisted of three or four shafts squashed together and connected with tight, 180 degree bends. Members of the genus Hamites were much larger but had a similar sort of shape, though the shafts were open so that the whole thing looked rather like a big paper clip.

The Late Cretaceous enjoyed the widest variety of heteromorphs, including the straight-shelled Baculites and Sciponoceras ; the helically coiled Turrilites ; and the bizarre Nipponites , which looked more like a ball of string than anything else. Some forms combined different coiling modes. For example, Nostoceras started off with a helix like a Turrilites, but had a planar hook hanging underneath, more like an Ancyloceras.

Not all the Ancyloceratida had such strange shaped shells. Many of the earlier forms had regularly coiled shells barely distinguishable from the regular ammonites, for example, the Lower Cretaceous genus Deshayesites . Some offshoots of the uncoiled varieties even went back to being regularly coiled. Most notable among these were the Scaphites, Hoploscaphites , and their relatives, which were mostly regular spirals in shape except for a very slight hook at the end.

The biology of the heteromorph ammonites is not clear, but one certainty is that the uncoiled shells would have made these forms very poor swimmers. Open shells, particularly ones with spines and ribs, create lots of drag; but more importantly, the orientation of the shell, with the body hanging below the buoyant part of the shell, would have created a serious impediment to efficient swimming. More likely these ammonites either drifted in the plankton, collecting small animals on long tentacles like modern jellyfish, or else they crawled along the sea floor feeding on sessile or slow-moving animals such as clams.

Ancyloceratida varied widely in size, ranging from diminutive Ptychoceras barely 3 cm long to Baculites and Diplomoceras that can be 1-2 m in length. Some species were very widely distributed, for example some species of Hamites can be found in Eurasia, South America, Australia, and Antarctica. Others, like those inhabiting the Western Interior Seaway that covered much of the US, were much more localized.

See also

• Nautiloid
• Lituites
• Orthoceras

External links

• Photos of heteromorph ammonites from the UK
• Diagrams of what heteromorphs may have looked like in life